School of Biotechnology and Biomolecular Sciences, University of New South Wales, Sydney, Australia
      Home Page :: Bill Ballard

Current Position

Professor (2006- current)
Biotechnology and Biomolecular Sciences
University of New South Wales.
Sydney, Australia

Previous Positions

Head of School (2008-2013)
Biotechnology and Biomolecular Sciences.
University of New South Wales,
Sydney, Australia

Professor (2001-2006)
Biology, University of Iowa
Iowa City, Iowa.

Pritzker Curator (1998-2001)
The Field Museum, Chicago, IL.

Assistant Curator (1995-1998)
The Field Museum, Chicago, IL.

Tel: 61+ 2 9385 2029


Postdoctoral Fellow (1992-1995)
NHMRC C.J. Martin Postdoctoral Fellow. University of Chicago, IL.

Postdoctoral Fellow (1990-1992)
CSIRO Division of Entomology, Canberra, Australia.

Ph.D (1986-1990) 
University of Queensland, Brisbane, Australia.

M.Sc./D.I.C. (1984-1985)
Imperial College, London, England.

B.Sc. (Hons) (1979-1983)
Australian National University, Canberra, Australia.


Towarnicki, G.T. and J.W.O. Ballard, Accepted 7/6/2017. Drosophila mitotypes determine developmental time in a diet and temperature dependent manner. J. Insect Physiol.

Melvin, R.G. and J.W.O. Ballard. 2017. Cellular and population level processes influence the rate, accumulation and observed frequency of inherited and somatic mtDNA mutations. Mutagenesis 32: 323-334

Aw, W., N.A Youngson and J.W.O. Ballard. 2016. Can we alter dietary macronutrient compositions and alleviate mitochondrial disease? J. Rare Dis. Treat. 1: 31-37.

Hew, J., S.M. Solon-Biet, A.C. McMahon, K. Ruohonen, D. Raubenheimer, J.W.O. Ballard, D. G. Le Couteur, Z. Li, P.K.M. Maitz, Y. Wang. and S. J. Simpson. 2016.  The effects of dietary macronutrient balance on skin structure in male and female mice. Plos One 11: e0166175

Bajracharya, R. and J.W.O. Ballard. 2016. Low protein to carbohydrate ratio diet delays onset of Parkinsonism like phenotype in Drosophila melanogaster parkin null mutants. Mech. Ageing Dev. 160: 19-27

Arendt, M., K. Cairns, J.W.O. Ballard, P. Savolainen and E. Axelsson. 2016. Diet adaptation in dog reflects spread of prehistoric agriculture. Heredity 117: 301-306.

Aw, W., S. Towarnicki, R. Bajracharya and J. W.O. Ballard. 2016. Assessing bioenergetic functions from isolated mitochondria in the fly Drosophila melanogaster. J. Biol. Methods e42. doi: 10.14440/jbm.2016.112

Castellano-Gonzalez, G., N. Pichaud, J.W.O. Ballard, A. Bessede, H. Marcal., G.J. Guillemin. 2016. Epigallocatechin-3-gallate induces oxidative phosphorylation by activating cytochrome c oxidase in human cultured neurons and astrocytes. Oncotarget 7: 7426-7440.

Ballard, J. W. O. and N.A Youngson. 2015. Can diet influence the selective advantage of mitochondrial DNA haplotypes? Biosci Rep. 35: e00277

Solon-Biet, S.M., K.A. Walters, U.K. Simanainen, A.C. McMahon, K. Rouhonen, J.W.O. Ballard, D. Raubenheimer, D.J. Handelsman, D.G. Le Couteur, and S.J. Simpson. 2015. Macronutrient balance, reproductive function, and lifespan in aging mice. Proc. Natl. Acad. Sci USA 112: 3481-6.

Le Couteur, D.G., S.S. Tay, S.M. Solon-Biet, P. Bertolino, A.C. McMahon, V.C. Cogger, F. Colakoglu, A. Warren, A. J. Holmes, N. Pichaud, M. Horan, C. Correa, R. G. Melvin, N. Turner,  J.W.O. Ballard, K. Ruohonen, D. Raubenheimer and S.J. Simpson. 2014. The influence of macronutrients on splanchnic and hepatic lymphocytes in aging mice. J. Gerontol. Biol Sci. doi:10.1093/gerona/glu196

Song, W-H, J.W.O. Ballard , Y-J Yi and P. Sutovsky. 2014. Regulation of mitochondrial genome inheritance by autophagy and ubiquitin-proteasome system: Implications for health, fitness, and fertility. BioMed Res. Inter. doi:10.1155/2014/981867

Chung, P.P., I. Chu and J.W.O. Ballard. 2014. Assessment of temporal genetic variability of two epibenthic amphipod species in an eastern Australian estuarine environment, and their suitability as biological monitors. Aust. J. Zool. 62:206-15

Solon-Biet, S.M., A.C. McMahon, J.W.O. Ballard, K. Ruohonen, L.E. Wu, V.C. Cogger, A. Warren, N. Pichaud, R.G. Melvin, R. Gokarn, M. Khalil, N. Turner, G.J. Cooney, D.A. Sinclair, D. Raubenheimer, D.G. Le Couteur, and S.J. Simpson. 2014. The ratio of macronutrients, not caloric intake, dictates cardiometabolic health, aging and longevity in ad libitum-fed mice. Cell Metab. 19: 418-430

Correa, C.C. and J.W.O. Ballard  2014. What can symbiont titres tell us about co-evolution of Wolbachia and their host? J. Invert. Pathol. 118: 20-27

Braidy, N., W-P Gai, Y.H. Xu, P. Sachdev, G.J. Guillemin, X-M Jiang, J. W.O. Ballard, M.P. Horan, Z.M. Fang, B.H. Chong, and D.K.Y. Chan. 2014. Alpha-synuclein transmission and mitochondrial toxicity in primary human foetal enteric neurons in vitro. Neurotox. Res. 25: 170-182

Cogger, V.C., D. Svistounov, A. Warren, S. Zykova, R. G. Melvin, S.M. Solon-Biet, J.N. O’Reilly, A.C. McMahon, J.W.O. Ballard, R. De Cabo D. G. Le Couteur and M. Lebel. 2014. Liver aging and pseudocapillarization in a Werner syndrome mouse model. J. Gerontol. A Biol. Sci. Med. Sci. PMID: 24149428

Ballard, J. W. O. and N. Pichaud. 2014. Mitochondrial DNA: More than an evolutionary bystander. Funct. Ecol. 28: 218-231

Braidy, N., W-P Gai, Y.H. Xu, P. Sachdev, G.J. Guillemin, X-M Jiang, J. W.O. Ballard, M.P. Horan, Z.M. Fang, B.H. Chong, and D.K.Y. Chan. 2013. Uptake and mitochondrial dysfunction of alpha-synuclein in human astrocytes, cortical neurons and fibroblasts. Trans. Neurodegen. 2: 20 doi: 10.1186/2047-9158-2-20

Aw, W. and J. W.O. Ballard. 2013. The effects of temperature and diet on age grading and population age structure determination in Drosophila.  J. Insect Physiol. 59: 994-1000.

Chung, P.P., J.W.O Ballard and R.V Hyne. 2013. Differential survival and reproductive performance across three mitochondrial lineages in Melita plumulosa following naphthalene exposure Chemospere 93: 1064-9

Chung, P.P., R.V Hyne, R. M Mann and J.W.O Ballard. 2013. The impact of historic isolation on the population biogeography of Melita plumulosa (Crustacea: Melitidae) in eastern Australia Est. Coast. Shelf Sci. 129: 198-205

Pichaud, N., M. Messmer, C. C. Corea and J. W. O. Ballard. 2013. Diet influences the intake target and mitochondrial functions of Drosophila melanogaster males. Mitochondrion 3: 817-822

Wolff, J.N., P. Sutovsky and J.W.O. Ballard. 2013. Mitochondrial DNA content of mature sperm and oocytes in the genetic model Drosophila. Cell Tissue Res. 353: 195-200

Pichaud, N., Garratt, M., J.W.O. Ballard and R.C. Brooks. 2013. Physiological adaptations to reproduction II. Mitochondrial adjustments in livers of lactating mice. J. Exp. Biol. 216: 2889-2895

Horan, M.P., J.N. Rumbly, R.G. Melvin, D.G. Le Couteur and J.W.O. Ballard. 2013. Quaternary protein modelling to predict the function of DNA variation found in human mitochondrial cytochrome c oxidase. J. Hum. Genet. 58:127-134

Pichaud, N., J.W. O Ballard, R. M. Tanguay and P. U. Blier. 2013. Mitochondrial haplotype divergences affect specific temperature sensitivity of mitochondrial respiration. J. Bioenerget. Biomem. 45:25-35

Wolff, J.N., M. Nafisinia, P. Sutovsky and J.W.O. Ballard. 2013. Paternal transmission of mitochondrial DNA as an integral part of mitochondrial inheritance in metapopulations of Drosophila simulans. Heredity 110: 57-62

Correa, C.C.  and J.W.O. Ballard  2012. Wolbachia gonadal density in female and male Drosophila vary with laboratory adaptation and respond differently to physiological and environmental challenges. J. Invert. Pathol. 111: 197-204

Aw, W., F. Dowell and J. W.O. Ballard. 2012. Determination of the potential for near infra-red spectroscopy to resolve the species, gender, age and the presence of Wolbachia in laboratory reared Drosophila. G3 2: 1057-1065

Horan, M.P. and J.W.O. Ballard. 2012. Application of chromosome conformation capture (3c) to the study of human disease. In “eLS 2012”John Wiley & Sons Ltd, Chichester. DOI: 10.1002/9780470015902.a0024383

Correa, C.C.  W. C. Aw, R. G. Melvin and J. W. O. Ballard. 2012. Mitochondrial DNA variants influence mitochondrial bioenergetics in Drosophila melanogaster. Mitochondrion 12: 459-464

Solon, S., A. McMahon, J.W. O. Ballard, D. Raubenheimer, G.J. Cooney, D.G. Le Couteur and S.J. Simpson. 2012. The effects of balance of macronutrients on ageing and glucose tolerance. Australas. J. Ageing 31: 33-33.

Blier, P.U., N. Pichaud, J.W.O. Ballard and R.M. Tanguay. 2012. Divergent mitochondrial haplotypes confer clear adjustments in metabolic phenotypes. Integr. Comp. Biol. 52: E15-E15

Pichaud, N., J.W.O. Ballard, R.M. Tanguay and P. U. Blier. 2012. Naturally occurring mitochondrial DNA haplotypes exhibit metabolic differences: Insight into functional properties of mitochondria. Evolution 66: 3189-3197

Wolff, J.N., R. Brooks, D. Shearman and J.W.O. Ballard. 2012. Selective enrichment and sequencing of whole mitochondrial genomes in the presence of nuclear encoded mitochondrial pseudogenes (Numts). PLoS One 7: e37142

Faith, D.P., F. Köhler, L. Puslednik and J.W.O Ballard. 2012. Corroboration assessments and recent progress towards integrative systematics: a reply to Farris and Carpenter. Zootaxa 323:65-68

Horan, M.P., N. Pichaud and J. W.O. Ballard. 2012. Quantifying mitochondrial dysfunction in complex diseases of aging. J. Gerontol A Bio.l Sci. Med. Sci. 67: 1022-1035

Puslednik, L., R. C. Russell and J.W.O. Ballard. 2012. Phylogeography of the medically important mosquito Aedes (Ochlerotatus) vigilax (Diptera: Culicidae) in Australasia. J. Biogeo. 39: 1333-1346

Chung, P.P., R.V Hyne, R. M Mann and J.W.O Ballard. 2012. It’s all in the genes: how genotype can impact upon response to contaminant exposure and the implications for biomonitoring in aquatic systems. In “Environmental Contamination” (Ed J. K. Srivastava) Intech, Rijeka, Croatia Pp 111-136

Puslednik, L., D. K. Yeates, D. P Faith and J.W.O. Ballard. 2012. Protein- protein interactions and the cytochrome c oxidase DNA barcoding region. Syst. Entomol. 37: 229-236

Aw, W. C., C. C. Correa, D. J. Clancy and J. W. O. Ballard. 2011. Mitochondrial DNA variants in Drosophila melanogaster are expressed at the level of the organismal phenotype. Mitochondrion 11: 756-763

Pichaud, N., J.W.O. Ballard, R. Tanguay and P.U. Blier. 2011. Thermal sensitivity of mitochondrial functions in permeabilized muscle fibers from two populations of Drosophila simulans with divergent mitotypes. Am. J. Physiol. 301: R48-R59

Melvin, R.G. and J.W.O. Ballard. 2011. Females with a mutation in a nuclear encoded mitochondrial protein pay a higher cost of survival than do males in Drosophila. J. Gerontol. A Biol. Sci. Med. Sci. 66: 765-770

Faith, D.P., F. Köhler, L. Puslednik and J.W.O Ballard. 2011. Phylogenies with corroboration assessment. Zootaxa 2946: 52-56

Ballard, J. W. O. and R.G. Melvin . 2011. Early life benefits and later life costs of a two amino acid deletion in Drosophila simulans. Evolution 65: 1400-1412

Cairns, K.C., A. Wilton and J.W.O Ballard. 2011. The identification of dingoes in a background of hybrids. Advances in Genetics Research Series 6: 309-327. Nova Science Publishers, Inc.Hauppauge, NY

Chung, P.P., R.V Hyne, R. M Mann and J.W.O Ballard. 2011.Temporal and geographical genetic variation in the amphipod Melita plumulosa (Crustacea: Melitidae): Link of a localized change in haplotype frequencies to a chemical spill. Chemospere 82: 1050-1055

Chatelain, H., N. Pichaud, J. W. O. Ballard, R. M. Tanguay, G. Morrow and P. U. Blier 2011. Functional conservatism among Drosophila simulans flies experiencing different thermal regimes and mitochondrial DNA introgression. J. Exp. Zool. Part B: Ref: JEZ-B-2010-02-0010.R1

Ballard, J.W.O., R.G. Melvin, M. Lazarou, F.J. Clissold and S.J. Simpson. 2010. Cost of a naturally occurring two-amino acid deletion in cytochrome c oxidase subunit 7A in Drosophila simulans. Am. Nat. 176: E98-E108

Pichaud, N., E.H. Chatelain, J.W.O. Ballard, R. Tanguay, G. Morrow and P.U. Blier. 2010. Thermal sensitivity of mitochondrial metabolism in two distinct mitotypes of Drosophila simulans: Evaluation of mitochondrial plasticity. J. Evol. Biol. 213: 1666-1675

Ballard, J.W.O and R.G. Melvin. 2010. Linking the mitochondrial genotype to the organismal phenotype. Mol. Ecol. 19: 1523-1539

Cairns, K.C., J.N. Wolff, R.C. Brooks and J.W.O. Ballard. 2010. Evidence of recent population expansion in the field cricket Teleogryllus commodus. Aust. J. Zool. 58: 33-38.

Ballard, J.W.O., L. Puslednik, J.N. Wolf and R.C. Russell. 2009. Variation under nature: A sesquicentennial  DNA barcoding perspective. Chiang Mai J. Sci. 36: 188-199

Ballard, J.W.O., R.G. Melvin and S.J. Simpson. 2008. Starvation resistance is positively correlated with body lipid proportion in five wild caught Drosophila simulans populations. J. Insect Physiol. 54: 1371-1376

Chung, P.P., R.V Hyne, R. M Mann and J.W.O Ballard. 2008. Genetic and life-history trait variation of the amphipod Melita plumulosa from polluted and unpolluted waterways in eastern Australia. Sci. Tot. Envir. 403: 222-229

Van Voorhies, W.A., Melvin, R.G., J.W.O Ballard and J. B. Williams. 2008. Validation of manometric microrespirometers for measuring oxygen consumption in small arthropods. J. Insect Physiol. 54: 1132-1137

Lee, K.P., S.J. Simpson, F.J. Clissold, R. Brooks, J.W.O Ballard and D. Raubenheimer. 2008. Lifespan and reproduction in Drosophila: new insights from nutritional geometry. Proc. Nat. Acad. Sci. U.S.A.105: 2498-2503

Melvin, R.G., S.D. Katewa and J.W.O. Ballard. 2008. A candidate complex approach to study functional mitochondrial DNA changes: Sequence variation and quaternary structure modeling of Drosophila simulans cytochrome c oxidase. J. Mol. Evol. PMID: 18320260

Kenny, M.C., A. Wilton and J.W.O. Ballard. 2008. Seasonal trade-off between starvation resistance and cold resistance in temperate wild-caught Drosophila simulans. Aust. J. Entomol. 47: 20-23

Melvin, R.G., W.A. Van Voorhies and J.W.O. Ballard. 2007. Working harder to stay alive: Metabolic rate increases with age in Drosophila simulans but does not correlate with life span. J. Insect Physiol. 53: 1300-1306

Ballard, J.W.O., S.D. Katewa, R.G. Melvin and G. Chan. 2007. Comparative analysis of mitochondrial genotype and aging. Ann. NY Acad. Sci. 1114: 93-106

Ballard, J.W.O., R.G. Melvin, J. T. Miller and S.D. Katewa. 2007. Sex differences in survival and disparity of mitochondrial bioenergetics in Drosophila simulans. Aging Cell 6: 699-708

Ballard, J.W.O. and R.G. Melvin. 2007. Tetracycline treatment influences mitochondrial metabolism and mtDNA density two generations after treatment in Drosophila. Insect Mol. Biol. 16: 799-802

Ballard, J.W.O., R.G. Melvin, S.D. Katewa and K. Maas. 2007. Mitochondrial DNA variation is associated with measurable differences in life history traits and mitochondrial metabolism in Drosophila simulans. Evolution 61:1735-1747

Katewa, S.D. and J.W.O. Ballard. 2007b. Sympatric Drosophila simulans flies with distinct mtDNA show age related differences in mitochondrial metabolism. Insect Biochem. Mol. Biol. 37: 923-932

Katewa, S.D. and J.W.O. Ballard. 2007a. Sympatric Drosophila simulans flies with distinct mtDNA show difference in mitochondrial respiration and electron transport. Biochem. Mol. Biol. 37: 213-222.

Katewa, S.D., R.G. Melvin and J.W.O. Ballard. 2006. Comparison of methods and body parts for estimation of hydrogen peroxide production by isolated mitochondria from Drosophila simulans. Dros. Inf. Serv. 89:107-111

Melvin, R.G. and J.W.O. Ballard. 2006. Intraspecific variation in survival and mitochondrial oxidative phosphorylation in wild-caught Drosophila simulans. Ageing Cell 5: 225-233

Ballard, J.W.O. 2005. Mini Review: Drosophila simulans as a novel model for studying mitochondrial metabolism and aging. Exp. Gerentol. 40: 763-773

Ballard, J.W.O. and D. M. Rand. 2005. The population biology of mitochondrial DNA and its phylogenetic implications. Ann. Rev. Ecol. Evol. Syst. 36: 621-642

Dean, M.D. and J.W.O. Ballard. 2005. High divergence among Drosophila simulans mitochondrial haplogroups arose in midst of long term purifying selection. Mol. Phylo. Evol. 36: 328-337

Karr, T.L. and J.W.O. Ballard. 2005. “Wolbachia and speciation” Encylopedia of Sciences (Nature Publishing Group)

Dean, M.D. and J.W.O. Ballard. 2004. Linking phylogenetics with population genetics to reconstruct the geographic origin of a species. Mol. Phylo. Evol.32: 998-1009

Ballard, J.W.O. and A. C. James. 2004. Differential fitness of mitochondrial DNA in perturbation cage studies correlates with global abundance and population history in Drosophila simulans. Proc. R. Soc. of Lond. B 271: 1197-1201

Charlat, S., J.W.O. Ballard and H. Merçot. 2004. What maintains non cytoplasmic incompatibility inducing Wolbachia in their hosts: a field study in Drosophila yakuba. J. Evol. Biol. 17: 322-330

Ballard, J.W.O. and M. C. Whitlock. 2004. The incomplete natural history of mitochondria. Mol. Ecol. 13: 729-744

Ballard, J.W.O. 2004. Sequential evolution of a symbiont inferred from the host: Wolbachia and Drosophila simulans. Mol. Biol. Evol. 21: 428-442

Ballard, J.W.O. 2004. Genetics in the Australian Cattle Dog. 2004 Australian Cattle Dog Annual. Hoflin Publishing Colorado, USA

Dean, M. D., K. J. Ballard, A. Glass, and J.W.O. Ballard. 2003. Influence of two Wolbachia strains on population structure of east African Drosophila simulans. Genetics 165: 1959-1969

James, A.C. and J.W.O. Ballard. 2003. Mitochondrial genotype affects fitness in Drosophila simulans. Genetics 164:187-194

Ballard, J.W.O., B. Chernoff, and A. C. James. 2002. Divergence of mitochondrial DNA is not corroborated by nuclear DNA, morphology or behavior in Drosophila simulans. Evolution 56: 527-545

James, A.C., M.D. Dean, M.E. McMahon, and J.W.O. Ballard. 2002. Dynamics of double and single Wolbachia infections in Drosophila simulans from New Caledonia. Heredity 88: 182-189

Ballard, J.W.O. and M.D. Dean. 2001. The mitochondrial genome: mutation, selection and recombination. Curr. Opin. Genet. Devel. 11:667-672

Dean, M.D. and J.W.O. Ballard. 2001. Factors affecting mitochondrial DNA quality from museum preserved Drosophila simulans. Ent. Exp. Appl. 98: 279-283

Steinbachs, J.C., N.V. Schizas and J.W.O. Ballard. 2001. Efficiency of different genes and accuracy of different methods in recovering a known Drosophila genealogy. Proceedings of the Pacific Symposium on Biocomputing 2001, (eds R.B. Altman, A.K. Dunker, L. Hunter, K. Lauderdale and T. E. Klein). Pp 606-617. World Scientific, Singapore

James, A.C. and J.W.O. Ballard. 2000. The expression of cytoplasmic incompatibility and its impact on population frequencies and the distribution of Wolbachia strains in Drosophila simulans. Evolution 54: 1661-1672

Ballard, J.W.O. 2000c. When one is not enough: introgression in Drosophila. Mol. Biol. Evol. 17: 1126-1130

Ballard, J.W.O. 2000b. Comparative genomics of mitochondrial DNA in Drosophila simulans. J. Mol. Evol. 51: 64-75

Ballard, J.W.O. 2000a. Comparative genomics of mitochondrial DNA in members of the Drosophila melanogaster subgroup. J. Mol. Evol. 51: 48-63

Shuster, S.M., J.W.O. Ballard, G. Zinser, C. Sassaman, and P. Keim. 2000. The influence of genetic and extrachromosomal factors on population sex ratio in the marine isopod, Paracerceis sculpta (Holmes). Proceedings of the second international conference on Isopod biology, (eds R.C. Brusca and B. Kensley). Balkema Press, Amsterdam

Ballard, J.W.O., M.K. Thayer, A.F. Newton and E.R. Grismer. 1998. Data sets, partitions and characters: Philosophies and procedures for analyzing multiple data sets. Syst. Biol. 47: 367-396

Patterson, B.D., J.W.O. Ballard and R.L. Wenzel. 1998. Distributional evidence for cospeciation between neotropical bats and their bat fly extoparasites. Stud. Neotrop. Fauna & Environm. 33: 76-84

Ballard, J.W.O., J. Hatzidakis, T.L. Karr and M. Kreitman. 1996. Reduced variation in Drosophila simulans mitochondrial DNA. Genetics 144: 1519-1528

Ballard, J.W.O. 1995. Data on the Web. Science 270: 13-14

Ballard, J.W.O. and M. Kreitman, 1995. Is mitochondrial DNA a strictly neutral marker? TREE 10: 485-488

Ballard, J.W.O. and M. Kreitman. 1994. Unraveling selection in the mitochondrial genome of Drosophila. Genetics 138: 757-772

Ballard, J.W.O. 1998. Evolutionary rate variation within the mitochondrial DNA of Drosophila simulans. Current Topics of Molecular Evolution. Proceedings of the TriNational Workshop on Molecular Evolution, Munich, Germany, (eds M.K. Uyenoyama and A von Haeseler). Pp. 25-36. Duke University Publications Group, Duke University, Durham, North Carolina

Brust, R.A., J.W.O. Ballard, F. Driver, D.M. Hartley, N.J. Galway and J. Curran. 1998. Molecular systematics and hybrid crossing identify a third taxon, Aedes (Halaedes) wardangensis sp.n., of the Aedes (Halaedes) australis species group (Diptera: Culicidae). Can. J. Zool. 76:1236-1246

Ballard, J.W.O. 1996. Combining data in phylogenetic analysis. TREE 11: 334

Ballard, J.W.O. and N.J. Galway. 1996. Assessing evolutionary hypotheses generated from mitochondrial DNA: inferences from Drosophila. In: Floyd, R.B., A.W. Sheppard and P.J. DeBarro (eds) Frontiers of Population Ecology. CSIRO Publishing, Collingwood, Australia pp. 473-483

Ballard, J.W.O. 1994. Evidence from 12S ribosomal RNA resolves a morphological conundrum in Austrosimulium (Diptera: Simuliidae). J. Aust. Ent. Soc. 33: 131-135

Ballard, J.W.O. 1994. Phylogenetic affinities of Axinia (Diptera: Axiniidae) as inferred from 12S rRNA sequences pp. 528-534. Appendix to: D.H. Colless, "New family of muscoid Diptera, with sixteen new species in four new genera (Diptera: Axiniidae)." Invert. Zool. 8: 471-534

Curran, J., F. Driver, J.W.O. Ballard and R.J. Milner. 1994. Phylogeny of Metarhyzium: Sequence analysis of the internal transcribed spacer and 5.8S region of the ribosomal DNA repeat. Mycol. Res. 98: 547-552

Faith, D.P. and J.W.O. Ballard. 1994. Length differences and topology-dependent tests: A response to Kallersjo et al. Cladistics 10: 57-64

Elder, R.J. and J.W.O. Ballard. 1993. Blackflies and the Rockhampton floods. Bull. Mosq. Cont. Assoc. Aust. 5:11-19

Ballard, J.W.O. and R.J. Elder. 1992. Monitoring blackfly (Diptera: Simuliidae) pests. J. Aust. Ent. Soc. 31: 263-270

Ballard, J.W.O., G.J. Olsen, D.P. Faith, W.A. Odgers, D.M. Rowell, and P.W. Atkinson. 1992. Evidence from 12S ribosomal RNA that onychophorans are modified arthropods. Science 258: 1345-1348

Ballard, J.W.O. 1991. Colonization of perspex strips by larvae of Austrosimulium bancrofti (Taylor)(Diptera: Simuliidae), near Willawarin, New South Wales. Aust. J. Zool. 39: 201-210

Ballard, J.W.O. and D.G. Bedo. 1991. Population cytogenetics of Austrosimulium bancrofti (Taylor) (Diptera: Simuliidae) in eastern Australia. Genome 34: 338-353

Ballard, J.W.O., M.P. Zalucki. and K.E. Basford. 1991. Colonization of perspex strips by larvae of Austrosimulium bancrofti (Taylor)(Diptera: Simuliidae), near Ipswich, Queensland. Hydrobiologia 218: 255-263

Ballard, J.W.O. and K.J. Morton. 1990. Populational differences in the trap- and host-finding behavior of Austrosimulium bancrofti (Taylor) (Diptera: Simuliidae) in eastern Australia. Can. J. Zool. 68: 579-584

Ballard, J.W.O. 1989. Factors influencing silhouette trap captures of the blackfly Austrosimulium bancrofti (Taylor) (Diptera: Simuliidae) in the Australian Capital Territory. Bull. Ent. Res. 79: 421-428

Ballard, J.W.O. 1988. A simple technique for sexing blackfly larvae of the taxon Austrosimulium bancrofti. Trans. R. Soc. Trop. Med. Hyg. 82: 478

Ballard, J.W.O. and A. Barnes. 1988. Factors influencing silhouette trap captures of the blackfly Austrosimulium bancrofti in Queensland, Australia. Med. Vet. Entomol. 2: 371-378

Ballard, J.W.O. and J.K. Waage. 1988. Feeding strategies of the horseflies Hybomitra expollicata and Tabanus bromius in southern France. Med. Vet. Entomol. 2: 265-270

Ballard, J.W.O. and I.D. Marshall. 1986. An investigation of the potential of Aedes camptorhynchus (Thom.) as a vector of Ross River virus. Aust. J. Exp. Biol. Med. Sci. 64: 197-200